In fact, we suggest that the tympanum of modern reptiles originated in the common ancestor of Parapleurota in the middle Permian. Excitingly, recent developmental work by Bronzati et al. has shown that the modern reptile tympanum was present in their common ancestor, congruent with this hypothesis

And I named one of them!!! Meet Aulacephalodon kapoliwacela - the Iron Hog - the first and so far only geikiid known from the Luangwa Basin! I worked on this taxon for my Master's thesis, and it's been lovely

When including remains described in 1977 by A. Perle (which unfortunately could not be located for the 2025 publication), we get a reasonable picture of much of this dinosaur’s skull.

Khankhuuluu is somewhat ‘transitional’ in both size and anatomy between the earlier tyrannosauroids and later, more familiar, tyrannosaurids like Tyrannosaurus itself.